The Grünbach Formation is an Austrian geological formation that dates to the early Campanian age of the Late Cretaceous. it forms part of the Gosau Group, and represents a marine regression event, representing a coastal/brackish environment, being underlain by the marine carbonate Maiersdorf Formation and overlain by the deep marine siliciclastic Piesting Formation. The main lithology is clay, marl, siltstone and sandstone, with a minor conglomerate component. Coal seams have also been noted.[1][2] It is notable for its fossils including those of dinosaurs and plants.
Vertebrate paleofauna
All remains have been found at a single locality, which is a thin marl seam in the Konstantin mining tunnel, within the Felbering Mine in the Neue Welt area north west of Muthmannsdorf in Lower Austria.[3] The initial remains were discovered in 1859 after an ornithopod tooth was found in a piece of coal in a dump outside the mine by Professor Ferdinand Stoliczka, and the productive seam discovered thereafter. The first material was described by Emanuel Bunzel in 1871[4] and then additional material was described by Harry Seeley in 1881.[5] Due to mining activity in the area ceasing at the end of the 19th century, no additional remains have been recovered since.
Squamates
| Squamates of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images |
| Araeosaurus | A. gracilis | Muthmannsdorf | | Vertebra[5] | | |
| Crocodyliformes of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images |
| Doratodon | D. carcharidens | Muthmannsdorf | | "incomplete mandible, a fragmentary right maxilla, a parietal fragment, and isolated teeth"[6] | | |
| Eusuchia | Indeterminate | Muthmannsdorf | | "mandible fragment and some postcranial material" | | |
Choristoderes
| Choristoderes of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images |
| Choristodera | Indeterminate | Muthmannsdorf | | "Two platycoelous vertebral centra"[7] | | |
Turtles
| Turtles of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images |
| Dortokidae[8] | Indeterminate | Muthmannsdorf | | Shell fragments | | |
| cf. Kallokibotioninae | Indeterminate | Muthmannsdorf | | Shell fragments | | |
Pterosaurs
| Pterosaurs of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images |
| "Ornithocheirus" | "O". buenzeli | Muthmannsdorf | | | Remains found not diagnostic, remains referred to Azdarchidae and Pterodactyloidea indet. | |
| Azdarchidae | Indeterminate | Muthmannsdorf | | "Proximal portion of a humerus"[9] | Previously referred to "O". buenzeli | |
| Pterodactyloidea | Indeterminate | Muthmannsdorf | | "Articular region of a lower jaw, crushed phalangeal fragments" | Previously referred to "O". buenzeli | |
Non-avian dinosaurs
| Dinosaurs of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images |
| Mochlodon | M. suessi | Muthmannsdorf | | Cranial and postcranial elements | | |
| Struthiosaurus | S. austriacus | Muthmannsdorf | | Cranial and postcranial elements | | |
| Tetanurae | Indeterminate | Muthmannsdorf | | "Two fragmentary teeth" | Referred to by the dubious name ‘Megalosaurus pannoniensis’[5] Nearly identical to teeth known from the Csehbánya Formation[10] | |
Flora
Most of these specimens were recovered from mining dumps near Grünbach am Schneeberg in lower Austria. The flora of the formation is considered to represent that of a high humidity subtropical climate, typical of the Euro-Sinian phytogeographical region.[11][12][13]
| | | |
Color key | | Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited. |
Bryophytes
| Bryophytes of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
| Stereisporites | Indeterminate | | | | Spore, representative of Sphagnaceae | |
Lycopodiophyta
| Lycopodiophytes of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
| Camarozonosporites | Indeterminate | | | | Spore, representative of Lycopodiaceae | |
| Retitriletes | Indeterminate | | | | Spore, representative of Lycopodium | |
Pteridophytes
| Pteridophytes of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
| Appendicisporites | A. tricuspidatus | | | | Spore, probable affinities with Schizaeaceae | |
| Cicatricosisporites | Indeterminate | | | | Spore, probable affinities with Schizaeaceae | |
| Cingulatisporites | Indeterminate | | | | Spore, affinities with Anemia | |
| Cladophlebis | C. gosauensis | | | | | |
| Coniopteris | ? | | | | | |
| Cyathidites | Indeterminate | | | | Spore | |
| Echinatisporites | Indeterminate | | | | Spore | |
| Equisetites | Indeterminate | | | | A horsetail | |
| Gosauopteris | G. danaeoides | | | | Probable affinities with Marattiaceae | |
| Gleicheniidites | G. senonicus | | | | Spore | |
| Leiotriletes | Indeterminate | | | | Spore, representative of Lygodium | |
| Marsileaceaephyllum | M. campanica | | | | Marsileaceae, very similar to extant Marsilea | |
| Matonisporites | Indeterminate | | | | Spore, Found in situ with Monheimia | |
| Microphyllopteris | M. austriaca | | | | | |
| Microtaenia | M. austriaca | | | | | |
| Monheimia | M. ungerii | | | | affinities with Matoniaceae, particularly Phanerosorus | |
| Polypodiaceoisporites | Indeterminate | | | | Spore | |
| Raphaelia | R. lobifolia | | | | | |
| Sphenopteris | S. gruenbachiana, S. ungeri | | | | Probable affinities with Schizaeaceae | |
Gymnosperms
| Gymnosperms of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
| Geinitzia | G. formosa, G. reichenbachii | | | | Pinopsida, Taxodioid affinity | |
| cf. Lindleycladus | L. lanceolatus | | | | | |
| Nilsonia | N. cf. holyi | | | | Bennettitales | |
| Pagiophyllum | Indeterminate | | | | | |
| Pityosporites | Indeterminate | | | | Pollen | |
| Podozamites | Indeterminate | | | | | |
| Taxiodiaceaepollenites | Indeterminate | | | | Pollen, at least 2 taxa | |
Angiosperms
Monocots
| Monocots of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
| Arecipites | Indeterminate | | | | Pollen, representative of Palmae | |
| Lysichiton | L. austriacus | | | | Araceae, genus extant | |
| Gruenbachia | G. pandanoides | | | | Pandanaceae, reproductive organs of Pandanites | |
| Pandanites | P. trinervis | | | | Pandanaceae | |
| Sabalites | S. longirhachis | | | | Palm | |
| Spinizonocolpites | Indeterminate | | | | Pollen, representative of extant Nypa palm | |
| Theiaiphyllum | T. kollmannii | | | | | |
Dicots
| Dicots of the Grünbach Formation |
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
| Brasenites | B. krasserii | | | | affinity with Brasenia | |
| Celastrophyllum | Indeterminate, C. johannae | | | | Hamamelidae | |
| Ceratoxylon | Indeterminate | | | | Reproductive structure | |
| Clavatipollenites | Indeterminate | | | | Pollen, representative of Chloranthaceae | |
| Complexiopollis | Indeterminate | | | | Pollen, affinities with Myricaceae | |
| Compositiphyllum | C. serratum | | | | | |
| Debeya | D. insignis | | | | | |
| Dicotylophyllum | Indeterminate | | | | | |
| Ettingshausenia | E. gruenbachiana, E. cf. laevis, Indeterminate | | | | Platanaceae | |
| Extremipollis | Indeterminate | | | | Pollen | |
| Grebenkia | G. europeica | | | | | |
| cf. Grevilleophyllum | G. constans | | | | | |
| Hungaropollis | Indeterminate | | | | Pollen | |
| Juglandiphyllites | J. pelagicus | | | | possible affinity with Juglandaceae | |
| Krutzschipollites | Indeterminate | | | | Pollen | |
| Laudaypollis | Indeterminate | | | | Pollen | |
| Leguminosites | L. mucronata | | | | | |
| Longanulipollis | Indeterminate | | | | Pollen | |
| Menispermites | M. ettingshausenii, M. summesbergerii | | | | | |
| Momipetes | Indeterminate | | | | Pollen, similar to extant Engelhardtia | |
| Myricophyllum | M. cf. zenkeri, M. serratum | | | | Affinities with Myrica | |
| Myrtacaedites | Indeterminate | | | | Pollen, Representative of Myrtaceae | |
| Normapolles | Indeterminate | | | | Pollen, very abundant. possibly associated with Juglandaceae | |
| Oculipollis | O. sp, O. parvoculus O.zaklinskaiaiae | | | | Pollen | |
| cf. Pandemophyllum | Indeterminate, P. cf. proteoides | | | | | |
| Pseudoplicapollis | Indeterminate | | | | Pollen | |
| Quereuxia | Q. angulata | | | | Possible affinity with Trapa | |
| Rodgersia | Indeterminate | | | | | |
| Semioculipollis | Indeterminate | | | | Pollen | |
| Suemegipollis | Indeterminate | | | | Pollen | |
| Ternstroemites | T. (?) neueweltensis | | | | | |
| Tricolpopollenites | Indeterminate | | | | Pollen, At least 8 distinct taxa | |
| Viburniphyllum | V. ermannii, V. austriacum | | | | | |
References
- ^ Draganits, Erich; Wagreich, Michael; Hofer, Gerald; Hofmann, Christa-Charlotte; Reischenbacher, Doris; Grundtner, Marie-Louise; Neuhuber, Stephanie; Bottig, Magda (2011). "Stratigraphy and geochemical characterisation of Upper Cretaceous non-marine - marine cycles (Grünbach Formation, Gosau Group, Austria". Austrian Journal of Earth Sciences. 104 (2): 90–107.
- ^ Grundtner, Marie-Louise (2011). Sedimentologie und Sedimentpetrographie der Gosau-Gruppe bei Dreistetten-Muthmannsdorf (Niederösterreichische Kalkalpen) (Master's thesis). University of Vienna. pp. 21–35, 51–57, 71. doi:10.25365/thesis.17461.
- ^ Sachs, Sven; Hornung, Jahn J. (May 2006). "Juvenile ornithopod (Dinosauria: Rhabdodontidae) remains from the Upper Cretaceous (Lower Campanian, Gosau Group) of Muthmannsdorf (Lower Austria)". Geobios. 39 (3): 415–425. doi:10.1016/j.geobios.2005.01.003. ISSN 0016-6995.
- ^ Bunzel, Emanuel (1871). "Die Reptilfauna der Gosau-Formation in der Neuen Welt bei Wiener-Neustadt". Abhandlungen der Kaiserlich-königlichen Geologischen Reichsanstalt. 5: 1–18.
- ^ a b c Seeley, H. G. (1881-02-01). "The Reptile Fauna of the Gosau Formation preserved in the Geological Museum of the University of Vienna: With a Note on the Geological Horizon of the Fossils at Neue Welt, west of Wiener Neustadt, by Edw. Suess, Ph.D., F.M.G.S., &c., Professor of Geology in the University of Vienna, &c". Quarterly Journal of the Geological Society. 37 (1–4): 620–706. doi:10.1144/GSL.JGS.1881.037.01-04.49. ISSN 0370-291X. S2CID 219235284.
- ^ Buffetaut, Eric (1979). "Revision der Crocodylia (Reptilia) aus den Gosau-Schichten (Ober-Kreide) von Österreich". Beiträge zur Paläontologie von Österreich. 6: 89–105.
- ^ Buffetaut, Eric (1989). "Erster nachweis von Choristodera (Reptilia, Diapsida) in der Oberkreide Europas: Champsosaurierwirbel aus den Gosau-Schichten (Campan) Niederösterreichs". Sitzungsberichten der Österreichs Akademis der Wissenschaften Mathematisch-Naturwissenschaftlichen Klasse, Abteilung. 197: 389–394.
- ^ Rabi, Márton; Vremir, Mátyás; Tong, Haiyan (2012-09-01), "Preliminary Overview of Late Cretaceous Turtle Diversity in Eastern Central Europe (Austria, Hungary, and Romania)", Vertebrate Paleobiology and Paleoanthropology, Springer Netherlands, pp. 307–336, doi:10.1007/978-94-007-4309-0_19, ISBN 9789400743083
- ^ Buffetaut, Eric; Ősi, Attila; Prondvai, Edina (March 2011). "The pterosaurian remains from the Grünbach Formation (Campanian, Gosau Group) of Austria: a reappraisal of 'Ornithocheirus buenzeli'". Geological Magazine. 148 (2): 334–339. Bibcode:2011GeoM..148..334B. doi:10.1017/S0016756810000981. ISSN 1469-5081. S2CID 131376979.
- ^ Csiki-Sava, Zoltan; Buffetaut, Eric; Ősi, Attila; Pereda-Suberbiola, Xabier; Brusatte, Stephen L. (2015-01-08). "Island life in the Cretaceous - faunal composition, biogeography, evolution, and extinction of land-living vertebrates on the Late Cretaceous European archipelago". ZooKeys (469): 1–161. doi:10.3897/zookeys.469.8439. ISSN 1313-2970. PMC 4296572. PMID 25610343.
- ^ Herman, Alexei B.; KVAČEK, JIŘÍ (2007). "Early Campanian Grünbach flora of Austria: systematic composition and palaeoclimatic interpretations". Acta Palaeobotanica. 47 (1): 37–55.
- ^ KVACEK, JIRÍ; HERMAN, ALEXEI (2004). "The Campanian Grünbach Flora of Lower Austria: palaeoecological interpretations". Annalen des Naturhistorischen Museums in Wien. 106 A.
- ^ B., Herman, Alexei (2010). Late Cretaceous Grünbach Flora of Austria. Kvaček, Jiří. Wien: Naturhistorisches Museum. ISBN 9783902421432. OCLC 517260675.
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