The Dakhla Formation , also called the Dakhla Shale ,[ 5] Maastrichtian -Danian  geologic formation  in the Western Desert , Egypt .[ 4] [ 6] Dinosaur  remains are among the fossils that have been recovered from the formation, although none have yet been referred to a specific genus .[ 7] 
 
Fossil content                Color key    Notes small text ; crossed out  taxa are discredited.  
 
Reptiles  
Dinosaurs   
Turtles   
Mosasaurs     Mosasaurs  reported from the Dakhla Formation   Genus  Species  Presence   Material Notes  Images     Globidens G. phosphaticus  Beris Oyster Mudstone Member[ 2]   A single tooth crown (CUNV0011).[ 2]   A mosasaurine .      Halisaurus H. hebae  Beris Oyster Mudstone Member[ 11]      A halisaurine         Mosasaurinae  Indeterminate   Beris Oyster Mudstone Member[ 2]   A fragmentary small dentary  with a single tooth (CUNV0012).[ 2]   A mosasaurine .        Prognathodon P . sp. "Lower part"   Two tooth crowns, three cervical, nine dorsal, three caudal vertebrae, and ribs, found in association (NVP025).[ 5]   Most similar to P. overtoni , possibly a member of that species      
 
Fish   
Invertebrates  
Bivalves     Bivalves  reported from the Dakhla Formation   Genus  Species  Presence   Material Notes  Images     Apectoichnus  A. longissimus  Ammonite Hill Member[ 1]   Borings in Nypa [ 1]   Bivalve borings.        Teredolites T. clavatus  Ammonite Hill Member[ 1]   32 moderately preserved borings (TBT02-33) & two badly preserved specimens (TBT44-45) in Nypa [ 1]   Bivalve borings.       
 
Plants   
See also    
References    ^ a b c d e f g   El Hedeny, M.; Kassab, W.; Rashwan, M.; Abu El-Kheir, G.; AbdelGawad, M. (2020-07-01). "Bivalve borings in Maastrichtian fossilNypa fruits: Dakhla Formation, Bir Abu Minqar, South Western Desert, Egypt" . Ichnos . 28  (1): 24– 33. doi :10.1080/10420940.2020.1784158 . ISSN  1042-0940 . S2CID  225527064 . ^ a b c d e f   AbdelGawad, Mohamed K.; Abu El-Kheir, Gebely A.; Kassab, Walid G. (September 2021). "The youngest records of mosasaurid reptiles from the Upper Cretaceous of the South-Western Desert in Egypt" . Proceedings of the Geologists' Association . 132  (5): 556– 562. Bibcode :2021PrGA..132..556A . doi :10.1016/j.pgeola.2021.07.004 . ISSN  0016-7878 . ^ "UPPER DAKHLA FORMATION ( BEIDA SHALE MEMBER) AT G. DUWI, RED SEA, EGYPT: MINERALOGICAL AND GEOCHEMICAL ASPECTS | Faculty of Science" . www.aun.edu.eg . Retrieved 2022-11-23  .^ a b c d e   Anan, Tarek I.; Abd El-Wahed, Ahmed G. (2017-12-15). "The Maastrichtian-Danian Dakhla Formation, Eastern Desert, Egypt: Utilization in manufacturing lightweight aggregates" Applied Clay Science . 150 : 10– 15. Bibcode :2017ApCS..150...10A . doi :10.1016/j.clay.2017.08.027 . ISSN  0169-1317 . ^ a b   El-Kheir, Gebely A. Abu; Shaker, Ahmed A.; Street, Hallie P.; Longrich, Nicholas R.; Strougo, Amin; Asan, Anhar; AbdelGawad, Mohamed (9 November 2023). "A Prognathodontin Mosasaur from the Maastrichtian of the Dakhla Oasis, Western Desert, Egypt" . Fossil Studies . 1  (1): 60– 75. doi :10.3390/fossils1010007 ISSN  2813-6284 . ^ Tantawy, A. A.; Keller, G.; Adatte, T.; Stinnesbeck, W.; Kassab, A.; Schulte, P. (2001-12-01). "Maastrichtian to Paleocene depositional environment of the Dakhla Formation, Western Desert, Egypt: sedimentology, mineralogy, and integrated micro- and macrofossil biostratigraphies" Cretaceous Research . 22  (6): 795– 827. Bibcode :2001CrRes..22..795T . doi :10.1006/cres.2001.0291 . ISSN  0195-6671 . ^ Weishampel, et al. (2004). "Dinosaur distribution." Pp. 517–607.  ^ a b c   Rauhut, Oliver W M; Werner, Christa (January 1997). "First record of a Maastrichtian sauropod dinosaur from Egypt" . Palaeontolgia Africana . 34 : 63– 67. ^ a b c d e f g h i j k l m n o   de Lapparent, France; Werner, Christa (April 1998). "New Late Cretaceous turtles from the Western Desert, Egypt" . Annales de Paléontologie . 84  (2): 131– 214. Bibcode :1998AnPal..84..131D . doi :10.1016/S0753-3969(98)80005-0 . ^ a b   Abu El-Kheir, G. A.-M.; AbdelGawad, M. K.; Kassab, W. G. (2021). "First known gigantic sea turtle from the Maastrichtian deposits in Egypt" . Acta Palaeontologica Polonica . 66  (2): 349– 355. ^ Shaker, A. A.; Longrich, N. R.; Strougo, A.; Asan, A.; Bardet, N.; Mousa, M. K.; Tantawy, A. A.; Abu El-Kheir, G. A. (2023). "A new species of Halisaurus  (Mosasauridae: Halisaurinae) from the lower Maastrichtian (Upper Cretaceous) of the Western Desert, Egypt". Cretaceous Research . 154 . 105719. doi :10.1016/j.cretres.2023.105719 . ^ a b   Capasso, Luigi; Abu El-kheir, Gebely Abdelmaksoud; Mousa, Mohamed Kamel; Tantawy, Abdel Aziz (December 2021). "ANOMOEODUS AEGYPTICUS N. SP. (PISCES, †PYCNODONTIFORMES) FROM THE LATE CRETACEOUS OF THE DAKHLA FORMATION, WESTERN DESERT, EGYPT" . Thalassia Salentina  (43): 89– 104. ^ a b   Abu El-Kheir; Tantawy; Mousa; Wahba; Capasso (2022-12-02). "Diastemapycnodus tavernensis gen. et sp. nov. (Actinopterygii, †Pycnodontiformes) from the marine Maastrichtian (Late Cretaceous) of the Dakhla Formation, Western Desert, Egypt" Historical Biology: A Journal of Paleobiology . 34  (12): 2324– 2331. Bibcode :2022HBio...34.2324A . doi :10.1080/08912963.2021.2014482 . S2CID  245051336 . ^ a b   Greenfield, T. (2025). "No evidence for a giant, late-surviving Onchopristis : Comment on Capasso et al. (2024)". Journal of African Earth Sciences . 223 . 105541. doi :10.1016/j.jafrearsci.2025.105541 . ^ a b   Capasso, L.; Abdel Aziz, S.; Tantawy, A. A.; Mousa, M. K.; Wahba, D. G. A.; Abu El-Kheir, G. A. (2024). "The first described Onchopristis  Stromer, 1917, (Elasmobranchii: †Onchopristidae) from the Marine Maastrichtian of Dakhla Formation, Western Desert, Egypt". Journal of African Earth Sciences . 105415. doi :10.1016/j.jafrearsci.2024.105415 . ^ El-Soughier, Maher I.; Mehrotra, R.C.; Zhou, Zhi-Yan; Shi, Gong-Le (January 2011). "Nypa fruits and seeds from the Maastrichtian–Danian sediments of Bir Abu Minqar, South Western Desert, Egypt" . Palaeoworld . 20  (1): 75– 83. doi :10.1016/j.palwor.2010.09.016 . ISSN  1871-174X .    
Bibliography  Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. 861 pp. ISBN  0-520-24209-2 .